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Botanical berries include tomatoes, not strawberries

In botany, a berry is a fleshy fruit from one flower and one ovary: grapes, tomatoes, bananas, even cucumbers. Culinary berries such as strawberries and raspberries fail that test. True berries keep an indehiscent pericarp; drupes instead harden a stony endocarp around the seed.

Everyday English calls any small, juicy, pitless fruit a berry. Botanical language is stricter: a simple fruit with seeds and fleshy pulp produced from a single flower’s ovary, without a split line that opens at maturity. The pericarp has three named layers—exocarp, mesocarp, endocarp—though botanists apply those labels inconsistently. Some sources say a berry’s endocarp should stay under two millimeters thick; a thicker, woody endocarp marks a drupe such as a peach or olive.

Specialized berry forms get their own names. Citrus fruits are hesperidia: thick-rinded berries divided into juicy segments. Gourd-family fruits are pepos—hard outer rind, no internal septa—covering cucumbers and melons. Berries from an inferior ovary may be called epigynous or false berries because floral-tube tissue joins the ovary; bananas, cranberries, blueberries, and many cucurbits fall here. Aggregate fruits such as raspberries fuse many ovaries of one flower, so they are not botanical berries. Strawberries are accessory fruits: the fleshy part is receptacle, not ovary.

Fleshy fruits appear late in the fossil record. Early flowering plants bore dry fruits; berries and drupes show up near the end of the Cretaceous or start of the Paleogene, about sixty-six million years ago, as vertebrate seed dispersers gained importance. In the nightshade family, dry capsules seem ancestral, and berries evolved at least three separate times. Carl Linnaeus’s 1751 Philosophia Botanica already set bacca apart from drupe and pome, yet fruit classification remains unsettled among botanists today.

Source: Berry (botany)

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